STEPPE PHYTOCOENOSES OF UKRAINE: SPESIFICS AND HISTORY OF THE STUDY

Keywords: steppe phytocenoses, abiotic and biotic factors of steppe formation, classification, mode of use, history of study.

Abstract

The article describes the characteristics of steppe plant groups, substantiates their ecological role, gives the main stages and directions of studying steppes, indicates the role of abiotic and biotic factors in their formation, in particular, the mechanism of influence of ungulate grazing on steppe plant groups. It is noted that the steppe ecosystem was formed under the influence of wild herbivores – herds of wild horses, kulans and roe deer (now their function is performed by domestic livestock), which is a necessary condition for maintaining the steppe type of vegetation. During grazing, the accumulation of dead plant biomass decreases, and during trampling by animals, the integrity of the dead cover is disturbed, which ensures the germination of new shoots of cereals and other steppe species. It also removes the problem of mesophytization due to the retention of a much larger amount of moisture by the dead cover than is typical for the steppes. Excessive grazing, as well as its complete absence, negatively affects the state of steppe vegetation. The description of the current state of steppe biotopes on the territory of Ukraine is given, and the high degree of their anthropogenic transformation is emphasized. Information is given about the areas of the steppe, including virgin steppe, which are now preserved on the territory of Ukraine and are part of the objects of the nature reserve fund. The biological features of typical steppe xerophyte species are indicated, including wax coating and pubescence on the leaves and a long and branched root system. The article includes information on aspects of anthropogenic activity, which became the main reason for the decrease in the share of steppe phytocenoses in the vegetation structure of Ukraine. The need for their preservation and protection is substantiated, taking into account their importance in preserving the valuable steppe biodiversity and their critical state. One of the main directions of studying steppe phytocenoses is the classification of steppe vegetation, the conditions and features of its formation, as well as the mode of loads that ensures the normal functioning of steppe ecosystems. The works of a number of both domestic and foreign scientists were analyzed.

References

1. Alehin, V.V. (1925). Fitosotsiologiya (uchenie o rastitelnyih soobschestvah) i eyo poslednie uspehi u nas i na Zapade [Phytosociology (the study of plant communities) and its recent successes in our country and in the West]. Metodika geobotanicheskih issledovaniy. Moskva, Puchnina, 7–75 (in Russian).
2. Alehin, V.V. (1957). Geografiya rasteniy s osnovami botaniki [Geography of plants with the basics of botany]. Moskva, 519 (in Russian).
3. Alehin, V.V. (1986). Teoreticheskie problemyi fitotsenologii i stepevedeniya [Theoretical problems of phytocenology and steppe science]. Moskva, 97 (in Russian). 4. Antifeev, D. A. & Golovanova, G. Ya. (2021). Ot Orlovo do Askanii [From Orlovo to Askania]. Melitopolskiy kraevedcheskiy zhurnal, № 17, 43–50 (in Russian).
5. Belgard, A.L. (1971). Stepnoe lesovedenie [Steppe forestry]. Moskva, Lesnaya promyishlennost, 336 (in Russian).
6. Bilyk, H. I. (1957). Roslynnist zapovidnyka Mykhailivska tsilyna ta yii zminy pid vplyvom hospodarskoi diialnosti liudyny [Flora of the Mykhailivska virgin reserve and its changes under the influence of human economic activity]. Ukr. botan. zhurnal, № 4, 26-39 (in Ukrainian).
7. Bilyk, H. I. (1973). Luchni stepy [Meadow steppes]. Roslynnist URSR. Stepy, kamianysti vidslonennia, pisky. Kyiv, Naukova dumka, 33–84 (in Ukrainian).
8. Bilyk, H. I. & Tkachenko V.S. (1972). Suchasnyi stan roslynnoho pokryvu zapovidnyka Mykhailivska tsilyna na Sumshchyni [The current state of the plant cover of the Mykhailivska Virgin Nature Reserve in Sumy Oblast]. Ukr. botan. zhurnal, 29, № 6, 696–702 (in Ukrainian).
9. Bilyk, H.I. (1973). Riznotravno-typchakovo-kovylovi stepy [Variegated-grassy-stemmed steppes]. Roslynnist URSR. Stepy, kamianysti vidslonennia, pisky. Kyiv, Naukova dumka, 94–179 (in Ukrainian).
10. Bilyk, H.I. (1974). Roslynnist luk balok zapovidnyka «Mykhailivska tsilyna» na Sumshchyni [Flora of meadows and beams of the reserve «Mykhailivskaya virgin land» in Sumy Oblast]. Ukr. botan. zhurnal, № 2, 198–204 (in Ukrainian).
11. Bilyk, H.I. & Osychniuk, V.V. (1967). Suchasnyi stan roslynnosti pokryv Askaniiskoho tsilynnoho stepu [The current state of vegetation cover of the Askaniya virgin steppe]. Ukrainskyi botanichnyi zhurnal, 24(4), 44–49 (in Ukrainian).
12. Bilyk, H.I. & Panova, L.S. (1959). Roslynni kompleksy zapovidnyka Kamiani Mohyly [Plant complexes of Kamiani Mohyly reserve]. Ukrainskyi botanichnyi zhurnal, 16(6), 40–48 (in Ukrainian).
13. Bilyk, H.I. & Tkachenko, V.S. (1971). Roslynnyi pokryv absoliutnoho zapovidnoi dilianky Khomutovskoho stepu [Vegetation cover of the absolutely protected area of the Khomutovsky Steppe]. Ukrainskyi botanichnyi zhurnal, 28(3), 337–342 (in Ukrainian).
14. Bilyk, H.I. & Tkachenko, V.S. (1971). Roslynnyi pokryv Provalskoho stepu URSR [Vegetation cover of the Proval steppe of the Ukrainian SSR]. Ukrainskyi botanichnyi zhurnal, 28(4), 443–449 (in Ukrainian).
15. Bilyk, H.I. & Tkachenko, V.S. (1971). Roslynnyi pokryv Striltsivskoho stepu URSR [Vegetation cover of the Striltsiv steppe of the Ukrainian SSR]. Ukrainskyi botanichnyi zhurnal, 28(5), 613–617 (in Ukrainian).
16. Bilyk, H.I. & Tkachenko, V.S. (1973). Zminy roslynnoho pokryvu stepu Mykhailivska tsilyna na Sumshchyni zalezhno vid rezhymu zapovidnosti [Changes in the vegetation cover of the Mykhailivska virgin steppe in Sumy Oblast depending on the regime of the nature reserve]. Ukr. botan. zhurnal, 30(1), 89–95 (in Ukrainian). 17. Borovyk, L.P. (2008). Pryrodni ta antropohenni faktory demutatsii perelohiv na terytorii Striltsivskoho stepu (viddilennia Luhanskoho pryrodnoho zapovidnyka) [Natural and anthropogenic factors of fallow demutation in the territory of the Striltsiv steppe (department of the Luhansk nature reserve)]. Chornomorskyi botanichnyi zhurnal, 4(1), 98–106 (in Ukrainian).
18. Borovyk, L.P. (2014). Vydovyi sklad perelohovykh uhrupovan pochatkovykh stadii suktsesii na pivnichnomu skhodi Luhanskoi oblasti [Species composition of fallow communities of the initial stages of succession in the north-east of Luhansk region]. Visnyk Lvivskoho universytetu. Seriia biolohichna, 64, 137–146 (in Ukrainian).
19. Didukh, Ya. P. & Korotchenko, I.A. (1996). Stepova roslynnist pivdennoi chastyny Livoberezhnoho lisostepu Ukrainy. I. Klasy Festucetea vaginatae ta Helianthemo-Thymetea [Steppe flora of the southern part of the Left Bank forest- steppe of Ukraine. I. Classes Festucetea vaginatae and Heliantgemo-Thytea]. Ukrainskyi fitotsenotychnyi zbirnyk. Seriia A. Fitosotsiolohiia, 2, 56–63 (in Ukrainian).
20. Didukh, Ya. P., Tkachenko, V. S., Pliuta, P. H., Korotchenko, I. A. & Fitsailo, T.V. (1998). Porivnialna otsinka fitoriznomanitnosti zapovidnykh stepovykh ekosystem Ukrainy z metoiu optymizatsii rezhymiv yikh okhorony [Comparative assessment of the phytodiversity of protected steppe ecosystems of Ukraine with the aim of optimizing their protection regimes]. Kyiv, 75 (in Ukrainian).
21. Didukh, Ya.P. (1989). Florystychna klasyfikatsiia uhrupovan «hisopovoi» flory [Floristic classification of groups of «hyssop flora»]. Ukrainskyi botanichnyi zhurnal, 46(6), 16–21 (in Ukrainian).
22. Didukh, Ya.P., Borsukevych, L.M., Davydova, A.O., Dziuba, T.P., Dubyna, D.V., Yemelianova, S.M., Kolomiichuk, V.P., Kuzemko, A.A., Kucher, O.O., Moisiienko, I.I., Pashkevych, N.A., Fitsailo, T.V., Khodosovtsev, O.Ie., Tsarenko, P.M., Chusova, O.O., Shapoval, V.V. & Shyriaieva, D.V. (2020). Biotopy stepovoi zony Ukrainy [Biotope of the steppe zone] (Red. akademik NAN Ukrainy Ya.P. Didukh). Kyiv – Chernivtsi, DrukART, 392 (in Ukrainian).
23. Didukh, Ya.P. & Korotchenko, I.A. (1996). Stepova roslynnist pivdennoi chastyny Livoberezhnoho lisostepu Ukrainy. I. Klasy Festucetea vaginatae ta Helianthemo-Thymetea [Steppe vegetation of the southern part of the Left Bank forest-steppe of Ukraine. I. Festucetea vaginatae and Helianthemo-Thymetea classes]. Ukrain skyi fitotsenotychnyi zbirnyk. Seriia A. Fitosotsiolohiia, 56–63 (in Ukrainian).
24. Didukh, Ya.P. & Vasheniak, Yu.A. (2012). Stepova roslynnist Tsentralnoho Podillia [Steppe vegetation of the Central Podillia]. Ukr. botan. zhurn., 69(6), 789–817 (in Ukrainian). 49
25. Dubyna, D.V. & Dziuba, T.P. (2007). Tsenotychne riznomanittia halofitnoi roslynnosti Ukrainy ufitosozolohichnomu aspekti [Cenotic diversity of halophytic vegetation of Ukraine from the phytosozological aspect]. Visti biosfernoho zapovidnyka «Askaniia-Nova», 9, 21–31 (in Ukrainian).
26. Dubyna, D.V. & Movchan, Ya.I. (2013). Ekomerezha stepovoi zony Ukrainy: pryntsypy stvorennia, struktura, elementy [Eco-network of the steppe zone of Ukraine: principles of creation, structure, elements]. Kyiv, LAT&K, 2013, 409 (in Ukrainian).
27. Dubyna, D.V., Sheliah-Sosonko, Yu.R., Zhmud, O.I., Zhmud, M.Ie., Dvoretskyi, T.V., Dziuba, T.P. & Tymoshenko, P.A. (2003). Dunaiskyi biosfernyi zapovidnyk [Danube Biosphere Reserve]. Roslynnyi svit. Kyiv, 459 (in Ukrainian).
28. Fitoriznomanittia zapovidnykiv i natsionalnykh pryrodnykh parkiv Ukrainy [Phytodiversity of reserves and national natural parks of Ukraine] (2012). Ch.1. Biosferni zapovidnyky. Pryrodni zapovidnyky. Kolektyv avtoriv pid red. V.A. Onyshchenka, T.L. Andriienko. Kyiv, Fitosotsiotsentr.
29. Hetman, V.I. (2000). Ekolohichni (edafichni) chynnyky produktyvnosti i bioriznomanittia stepovykh landshaftnykh kompleksiv Mykhailivskoi tsilyny [Ecological (edaphic) factors of productivity and biodiversity of the steppe landscape complexes of Mykhailivska virgin land]. Problemy landshaftnoho riznomanittia Ukrainy: Zbirnyk naukovykh prats. Kyiv, 175–182 (in Ukrainian).
30. Hryhora, I.M. & Solomakha, V.A. (2005). Roslynnist Ukrainy (ekoloho-tsenotychnyi, florystychnyi ta heohrafichnyi narys) [Vegetation of Ukraine (ecological-cenotic, floristic and geographical outline)]. Kyiv, Fitosotsiotsentr, 2005, 452 (in Ukrainian).
31. Hrytsenko, V. V. (2007). Luchni stepy Kyivskoho plato: flora, roslynnist, populiatsii ridkisnykh vydiv ta okhorona [Meadow steppes of the Kyiv plateau: flora, vegetation, populations of rare species and protection]: avtoref. dys. na zdobuttia nauk. stupenia kand. biol. nauk: 03.00.05. Kyiv, 20 (in Ukrainian).
32. Hrytsenko, V. V. (2017). Fitoriznomanittia botaniko-heohrafichnoi dilianky «Stepy Ukrainy» u Natsionalnomu botanichnomu sadu im. M.M. Hryshka NAN Ukrainy [Phytodiversity of the botanical and geographical area «Steppes of Ukraine» in the National Botanical Garden named after M.M. Hryshka of the National Academy of Sciences of Ukraine]. Lisove i sadovo-parkove hospodarstvo, 12 (in Ukrainian).
33. Podpera, J. (1925). Versuch einer epiontologischen Gliederung des europäischen Waldes. 21
34. Keller, B.A. (1923). Rastitelnyiy mir russkih stepey, polupustyin i pustyin [The flora of the Russian steppes, semi-deserts and deserts. Ecological and phytosociological essays]. Ocherki ekologicheskie i fitosotsiologicheskie, 1, 183 (in Russian).
35. Kharkevych, S. S. (1956). Stepovyi zapovidnyk Mykhailivska tsilyna [Steppe reserve Mykhailivska virgin land]. Ukrainskyi botanichnyi zhurnal, 13(2), 58–67 (in Ukrainian).
36. Kleopov, Yu.D. (1929). Do pytannia pro klasyfikatsiiu stepiv pivdennoho skhodu Ukrainy [To the question of the classification of the steppes of southeastern Ukraine]. Visnyk Kyivskoho botanichnoho sadu. Kyiv, 10, 41–52 (in Ukrainian).
37. Kolomiichuk, V.P., Ostapko V.M. & Yarovyi S.S. (2012). Pryrodnyi zapovidnyk Ukrainskyi stepovyi [Ukrainian Steppe Nature Reserve]. Fitoriznomanittia zapovidnykiv i natsionalnykh pryrodnykh parkiv Ukrainy. Ch. 1. Biosferni zapovidnyky. Pryrodni zapovidnyky. Kyiv, Fitosotsiotsentr, 336–378 (in Ukrainian).
38. Konaikova, V.O. (2018). Biotopy pryrodnoho zapovidnyka «Ielanetskyi step» [Biotopes of the nature reserve «Yelanetsky Steppe»]. U zb.: Klasyfikatsiia roslynnosti ta biotopiv Ukrainy: materialy tretoi naukovo-teoretychnoi konferentsii (Kyiv, 19–21 kvitnia 2018 r.). Kyiv, 87–92 (in Ukrainian).
39. Kondratyuk, E.N., Burda R.I., Chuprina T.T. & Homyakov M.T. (1988). Luganskiy gosudarstvennyiy zapovednik [Luhansk State Reserve]. Kiev, Naukova dumka, 188 (in Russian).
40. Kondratyuk, E.N. & Chuprina, T.T. (1992). Kovyilnyie stepi Donbassa. Sovremennoe sostoyanie i perspektivyi vosstanovleniya [Feather-grass steppes of Donbass. Current state and prospects for recovery]. Kiev, Naukova dumka, 171 (in Russian).
41. Korzhinskiy, S. I. (1888). Severnaya granitsa chernozemno-stepnoy oblasti vostochnoy polosyi Evropeyskoy Rossii v botaniko-geograficheskom i pochvennom otnoshenii [The northern border of the chernozem-steppe region of the eastern strip of European Russia in botanical-geographical and soil terms]. Kazan, 18(5), 253 (in Russian).
42. Kryilov, P.N. (1927). Flora Zapadnoy Sibiri [Flora of Western Siberia]. Tomsk, 212 (in Russian).
43. Kuzemko, A.А. (2016). Classification of the class Molinio-Arrhenatheretea in the forest and foreststeppe zones of Ukraine. Phytocoenologia, 46(3), 241–256.
44. Kuzemko, A. A. (2011). Kontseptsiia asotsiatsii v suchasnii fitosotsiolohii [The concept of association in modern phytosociology]. Chornomorskyi botanichnyi zhurnal, 7(3), 215–229 (in Ukrainian).
45. Kuzemko, A.А. (2009). Dry grasslands on sandy soils in the forest and forest-steppe zones of the plains region of Ukraine: present state of syntaxonomy. Tuexenia, 29, 369–390.
46. Kuzemko, A., Dubyna, D., Dziuba, T., Moysienko, I., Vasheniak, Y. & Zakharova, M. (2015). Syntaxonomy of sandy and rocky grasslands of Ukraine: preliminary results of large-scaleanalysis. In: From Population Biology to Community Ecology. 12 th European Dry Grassland Meeting. Book of Abstracts (Mainz, Germany, 22–27 May 2015). Mainz-Trier, 24.
47. Lavrenko, E. M. (1971). Osnovnyie problemyi biogeotsenologii i zadachi biogeotsenologicheskih issledovaniy v SSSR [The main problems of biogeocenology and the tasks of biogeocenological research in the USSR]. Zhurnal obschey biologii., 32 (4), 395–408 (in Russian).
48. Lavrenko, E. M. (1980). Stepi. Rastitelnost Evropeyskoy chasti SSSR [Steppes. Vegetation of the European part of the USSR]. Leningrad, Izd-vo Nauka, 203–272 (in Russian).
49. Lavrenko, E.M. (1954). Stepi Evraziatskoy stepnoy oblasti, ih geografiya, dinamika i istoriya [Steppes of the Eurasian steppe region. Their geography, dynamics, history]. Voprosyi botaniki. Leningrad, Nauka, 155 – 191 (in Russian).
50. Lavrenko, E.M., Karamyisheva, Z.V. & Nikulina, R.I. (1991). Stepi Evrazii [Steppes of Eurasia]. Nauka, 146 (in Russian).
51. Lavrenko, Ye.M. & Zoz, I.H. (1928). Roslynnist tsilyny Mykhailivskoho kinnoho zavodu (kol. Kapnista) Sumskoi okruhy [Roslinnistst tsiliny Mikhailivsky film factory (kol. Kapnist) Sumy district]. Okhorona pamiatok pryrody na Ukraini. Kharkiv, 2, 3–16 (in Ukrainian).
52. Lyisenko, G. N. (2014). Stepnyie zapovedniki i absolyutno zapovednyiy rezhim: poiski kompromissa [Steppe reserves and absolutely reserved regime: the search for a compromise]. Stepnoy byulleten, 11–16 (in Russian).
53. Lyisenko, G. N. & Korotchenko, I. A. (2006). Sintaksonomicheskie izmeneniya rastitelnogo pokrova lugovoy stepi zapovednika «Mihaylovskaya tselina» (Sumskaya oblast, Ukraina) [Syntaxonomic changes in the vegetation cover of the meadow steppe of the reserve «Mikhailovskaya virgin» (Sumy region, Ukraine)]. Rastitelnost Rossii, 43–57 (in Russian).
54. Lysenko, H. (2007). Krytychnyi period avtohenetychnoi suktsesii stepovykh fitotsenostruktur yak vyiav protsesiv filotsenohenezu [The critical period of the autogenetic succession of steppe phytocenous structures as a manifestation of the processes of phyllocenogenesis]. Lviv, Visnyk Lvivskoho universytetu, Seriia biolohichna, 45, 85–90 (in Ukrainian).
55. Mala, Yu.I. (2016). Mezha mizh Lisostepom i Stepom: ekoloho-tsenotychna otsinka (na prykladi pravoberezhnoi Ukrainy) [The border between the forest-steppe and the steppe: an ecological and cenotic assessment (on the example of the right-bank Ukraine)]. Kyiv, Naukova dumka,165 (in Ukrainian).
56. Mordkovych, V. H. (1982). Stepnye ecosistemy [Steppe ecosystems]. Novosybyrsk, Nauka, 1982, 206 (in Ukrainian).
57. Onyshchenko, V.A. & Andriienko, T.L. (2012). Fitoriznomanittia zapovidnykiv ta natsionalnykh pryrodnykh parkiv Ukrainy [Phytodiversity of reserves and national natural parks of Ukraine]. Kyiv, Fitosotsiotsentr, 406 (in Ukrainian).
58. Osychniuk, V. V. (1973). Zmina produktyvnosti stepovykh fitotsenoziv zalezhno vid struktury roslynnoho pokryvu [Change in the productivity of steppe phytocenoses depending on the structure of the plant cover]. Ukr. botan. zhurn., 30(2), 204–211 (in Ukrainian).
59. Osychniuk, V. V. (1979). Deiaki osoblyvosti zapovidnoho rezhymu u viddilenniakh Ukrainskoho stepovoho zapovidnyka [Some features of the protected regime in the branches of the Ukrainian Steppe Reserve]. Ukr. botan. zhurn., 36(4), 347–352 (in Ukrainian).
60. Osyichnyuk, V. V. (1988). Flora Ukrainskogo stepnogo zapovednika (annotirovannyiy spisok sosudistyih rasteniy) [Flora of the Ukrainian Steppe Reserve (an annotated list of vascular plants)]. Moskva, 44 (in Russian).
61. Pachoskiy, I. K. (1917). Opisanie rastitelnosti Hersonskoy gubernii, ch.2. Stepi [Description of the vegetation of the Kherson province, part 2. Steppes]. Materialyi po issledovaniyu pochv i gruntov Hersonskoy gubernii, 13, 336 (in Russian).
62. Pachoskiy, I. K. (1923). Tselinnaya zapovednaya step Askaniya-Nova [Virgin protected steppe Askania-Nova]. Askaniya-Nova, stepnoy zapovednik Ukrainyi, Moskva (in Russian).
63. Pachoskiy, I. K. (1926). Nablyudeniya nad tselinnyim pokrovom v Askaniya-Nova v 1923 g [Observation of virgin cover in Askania-Nova]. Visti Derzhavnogo stepovogo zapovidnika «ChaplI», 3 (in Russian).
64. Pohrebniak, P.S. (1968). Osnovy lisovoi typolohii [Basics of forest typology]. Moskva, Kolos, 440 (in Ukrainian).
65. Panova, L.S. (1972). Rozpodil roslynnoho pokryvu zapovidnyka Kamiani Mohyly zalezhno vid reliefu [Distribution of the vegetation cover of the Kamiani Mohyla reserve depending on the topography]. Ukrainskyi botanichnyi zhurnal, 29(4), 468–475 (in Ukrainian).
66. Panova, L.S. (1983). Rozpodil roslynnoho pokryvu zapovidnyka Kamiani Mohyly [Distribution of vegetation cover of Kamiani Mohyla reserve]. Ukrainskyi botanichnyi zhurnal, 40(3), 70–80 (in Ukrainian).
67. Parakhonska, N.O. (1984). Zminy florystychnoho skladu Mykhailivskoi tsilyny v umovakh zapovidnosti [Changes in the floristic composition of Mykhailivska virgin land under the conditions of the reserve]. Ukr. botan. zhurn., 41(5), 13–16 (in Ukrainian).
68. Polchaninova, N.Yu. (2004). Vosstanovlenie stepnyih araneokompleksov na zapovednyih territoriyah s razlichnyim senokosnyim rezhimom [Restoration of steppe araneocomplexes in protected areas with different haymaking regimes]. Aktualnyie problemyi sohraneniya ustoychivosti zhivyih sistem. Mater. VIII Mezhdunar. nauchn. ekol. konf. Belgorod, 149–165 (in Russian).
69. Polchaninova, N.Yu. (2012). Izmenenie stepnyih araneokompleksov pod vozdeystviem rezhimnyih meropriyatiy v zapovednikah Ukrainyi i Rossii [Changes in steppe araneocomplexes under the influence of regime measures in the reserves of Ukraine and Russia]. Rezhimyi stepnyih osobo ohranyaemyih prirodnyih territoriy: Materialyi mezhdunarodnoy nauchno-prakticheskoy konferentsii, posvyaschennoy 130-letiyu so dnya rozhdeniya professora V.V. Alehina. Kursk, 156–159 (in Russian).
70. Prystupa, I.V. (2011). Osnovy heobotaniky ta fitotsenolohii: Navchalnyi posibnyk [Basics of geobotany and phytocenology: Study guide]. Zaporizhzhia, ZNU, 10 (in Ukrainian).
71. Rabotnov, T. A. (1978). Fitotsenologiya: Uchebnoe posobie dlya biologicheskih fakultetov vuzov [Phytocenology: Textbook for biological faculties of universities]. Izdatelstvo MGU, Moskva, 384 (in Russian).
72. Rodinka, O.S. (2014). Florystychni zminy u zapovidnyku «Mykhailivska tsilyna» ta yikh prychyny [Floristic changes in the reserve «Mykhailivska virgin» and their causes]. Pryrodnychi nauky, 11, 52–57 (in Ukrainian).
73. Rodinka, O.S. & Shevchenko, Yu.M. (2013). Vplyv bahatorichnoho rezhymnoho vykoshuvannia travostoiu u viddilenni «Mykhailivska tsilyna» Ukrainskoho stepovoho pryrodnoho zapovidnyka na stan zberezhennia ta dynamiku chyselnosti vydiv roslyn ta tvaryn, zanesenykh do «Chervonoi Knyhy Ukrainy» [The effect of long-term regular mowing of grass in the «Mykhailivskaya virgin» section of the Ukrainian Steppe Nature Reserve on the state of preservation and the dynamics of the number of plant and animal species listed in the «Red Book of Ukraine»]. News Biosphere Reserve «Askania Nova», 15, 26 (in Ukrainian).
74. Shalyt, M. (1928). Deiaki vidomosti za protses vidnovlennia stepovoi roslynnosty na perelohakh Derzhavnoho stepovoho zapovidnyka «Chapli» (kol. Askaniia-Nova) [Some information on the process of restoration of steppe vegetation on the fallows of the State Steppe Reserve “Chapli” (Askania-Nova district)]. Visti Derzh. Stepovoho zapovidnyka «Chapli», 7, 139–152 (in Ukrainian).
75. Shalyt, M. (1928). Velykyi Chapelskyi pid v Askanii-Nova ta yoho roslynnist roku 1927-1928 [Velikiy Chapelsky pod in Askania-Nova and its vegetation in 1927–1928.]. Visti Derzh. Stepovoho zapovidnyka «Chapli», 7, 165–199 (in Ukrainian).
76. Shapoval, V.V. (2012). Biosfernyi zapovidnyk «Askaniia-Nova imeni F.E. Falts-Feina» [Biosphere Reserve «Askania- Nova named after F.E. Falz-Fein»]. V kn.: Fitoriznomanittia zapovidnykiv i natsionalnykh pryrodnykh parkiv Ukrainy. Ch.1. Biosferni zapovidnyky. Pryrodni zapovidnyky. Kyiv, Fitosotsiotsentr, 13–30 (in Ukrainian).
77. Shennikov, A.P. (1964). Vvedenie v geobotaniku [Introduction to Geobotany]. Izdatelstvo Leningradskogo universiteta, Leningrad, 448 (in Russian).
78. Teetzmann Franz. (1845). Ueber die Südrussischen Steppen und über die darinim Taurischen Gouvernement belegenen Besitzungen des Herzogs von Anhalt-Köthen. Beiträgezur Kenntniss des Russischen Reiches und der angränzenden Länder Asiens. Elftes Bändchen. St. Petersburg, 3–137.
79. Tkachenko, V.S., Lysenko, H.M. (2005). Synfitoindykatsiia postpirohennykh zmin ekotopichnykh kharakterystyk luchnoho stepu «Mykhailivska tsilyna» na Sumshchyni (Ukraina) [Synphytoindication of post-pyrogenic changes in the ecotopic characteristics of the meadow steppe «Mykhailivska virgin» in Sumy Oblast (Ukraine)]. Ukr. botan. zhurn., 62(4), 468–482 (in Ukrainian).
80. Tkachenko, V.S., Parakhonska, N.O. & Sheremet, L.H. (1984). Dynamika struktury roslynnoho pokryvu zapovidnyku «Mykhailivska tsilyna» [The dynamics of the structure of the plant cover of the reserve «Mykhailivska virgin land»]. Ukr. botan. zhurn., 41(3), 71–74 (in Ukrainian).
81. Tkachenko, V. S. (2004). Fitotsenotychnyi monitorynh rezervatnykh suktsesii v Ukrainskomu stepovomu pryrodnomu zapovidnyku [Phytocenotic monitoring of reserve successions in the Ukrainian Steppe Nature Reserve]. Kyiv, Fitosotsiotsentr, 184 (in Ukrainian).
82. Tkachenko, V. S., Henov, A. P. & Lysenko, H. M. (2003). Strukturni zminy v roslynnomu pokryvizapovidnoho luchnoho stepu «Mykhailivska tsilyna» (Ukraina) za dannymy velykomasshtabnohokartuvannia u 2001 rotsi [Structural changes in the vegetation cover of the protected meadow steppe “Mykhailivska virgin” (Ukraine) according to the data of large-scale mapping in 2001]. Visti biosf. Zapov. «Askaniia-Nova», 5, 7–17 (in Ukrainian).
83. Tkachenko, V. S., Henov, A.P. & Lysenko, H.M. (1991). Struktura roslynnosti zapovidnoho stepu «Mykhailivska tsilyna » za danymy krupnomasshtabnoho kartuvannia v 1991 rotsi [Vegetation structure of the protected steppe “Mykhailivska virgin land” according to the data of large-scale mapping in 1991]. Ukr. botan. zhurn., 50(4), 5–15 (in Ukrainian).
84. Tkachenko, V.S. (2007). Vtraty enerhii stepovymy ekosystemamy za riznykh vydiv yikh ekspluatatsii ta enerhetychni osnovy stabilnosti stepu [Energy losses of steppe ecosystems under different types of their exploitation and energy bases of steppe stability]. Ukrainskyi fitotsenotychnyi zbirnyk. Seriia S. Fitoekolohiia, 25, 4–18 (in Ukrainian).
85. Tkachenko, V.S. (2009). «Striltsivskyi step» v fitotsenotychnomu monitorynhu Starobilskykh stepiv [«Striltsivsky steppe» in the phytocenotic monitoring of the Starobil steppes]. Visti Biosfernoho zapovidnyka «Askaniia-Nova», 11, 6–19 (in Ukrainian).
86. Tkachenko, V.S., Didukh, Ya.P., Henov, A.P. & Dudka, I.O. (1998). Ukrainskyi pryrodnyi stepovyi zapovidnyk. Roslynnyi svit [Ukrainian natural steppe reserve. The plant world]. Kyiv, Fitosotsiotsentr, 280 (in Ukrainian).
87. Tkachenko, V.S. & Henov, A.P. (1986). Florotsenotychna kharakterystyka zaproponovanoho Kalmiuskoho derzhavnoho zakaznyka [Phylocoenotic characteristics of the proposed Kalmius state reserve]. Ukrainskyi botanichnyi zhurnal, 43(5), 92–96 (in Ukrainian).
88. Tymoshenko, P.A. (1999). Suchasnyi stan klasyfikatsii roslynnosti aren pivdnia Ukrainy [The current state of the vegetation classification of the arenas of southern Ukraine]. Ukrainskyi fitotsenotychnyi zbirnyk. Seriia A. Fitosotsiolohiia, 1–2(12–13), 201–205 (in Ukrainian).
89. Ukrainska radianska entsyklopediia u 12 tomakh. (1985). Stepova zona. Holovna redaktsiia URE [Ukrainian Soviet encyclopedia. Steppe zone] (in Ukrainian).
90. Vakarenko, L.P. (2009). Perspektivyi sozdaniya stepnyih regionalnyih parkov v Odesskoy oblasti [Prospects for the creation of steppe regional parks in the Odessa region]. Stepnoy byulleten, 26, 15–18 (in Russian).
91. Vasilyuk, A.V., Parnikoza, I.Yu. & Shevchenko, M.S. (2010). Bioraznoobrazie stepey pod ohranoy Krasnoy i Zelenoy knig Ukrainyi [The biodiversity of the steppes under protection of the Red and Green Books of Ukraine]. Stepnoy byulleten, №29, 33–38 (in Russian).
92. Vasyliuk, O. & Kaliuzhna, M. (2009). Lisorozvedennia u stepovii zoni Ukrainy: realii, obmezhennia, zahrozy [Forestry in the steppe zone of Ukraine: realities, limitations, threats]. Ekolohiia. Pravo. Liudyna, №4–5 (34–34), 35–48 (in Ukrainian).
93. Vasyliuk, O. & Kolomytsev, H. (2011). Implementatsiia Kiotskoho protokolu zahrozhuie stepam Ukrainy [The implementation of the Kyoto Protocol threatens the steppes of Ukraine]. Ekolohiia. Pravo. Liudyna, №13–14 (53–54), 29–31 (in Ukrainian).
94. Vedenkov, E. P. (1997). O vosstanovlenii estestvennoy rastitelnosti na yuge stepnoy Ukrainyi [On the restoration of natural vegetation in the south of the steppe Ukraine]. Askaniya-Nova, 39 (in Russian).
95. Vedenkov, E. P. & Drogobyich, N. E. (1998). O vosstanovitelnoy suktsessii zalezhnoy rastitelnosti v Askanii-Nova. Zapovidna sprava v Ukrayins [On the restorative succession of fallow vegetation in Askania-Nova]. Kaniv, t.3, vip.2, 81–85 (in Russian).
96. Voitiuk, B.Iu. (2005). Roslynnist zasolenykh gruntiv Pivnichno-Zakhidnoho Prychor-nomoria (suchasnyi stan, klasyfikatsiia, napriamky transformatsii, okhorona) [Vegetation of the saline soils of the North-Western Prychor- Nomor region (current state, classification, directions of transformation, protection)]. Kyiv, Fitosotsiotsentr, 224 (in Ukrainian).
97. Vysotskiy, G.M. (1905). Stepi Evropeyskoy Rossii [Steppes of European Russia]. Polnaya entsiklopediya russkogo selskogo hozyaystva, Moskva, 397–443 (in Russian).
98. Yakubenko, B. Ye., Hryhora, I. M. & Nikonov, S. B. (2005). Stepova roslynnist Ukrainy: suchasnyi stan ta perspektyvy yii optymizatsii ta vykorystannia [Steppe vegetation of Ukraine: current state and prospects for its optimization and use]. Monohrafiia: Naukove zabezpechennia staloho rozvytku silskoho hospodarstva v Stepu Ukrainy i AR Krym. Kabinet Ministriv Ukrainy. Natsionalnyi ahrarnyi universytet. Kyiv, Alefa, 1, 408–432 (in Ukrainian).
99. Yakubenko, B.Ie., Hryhoriuk, I.P., Yakubenko, N.B. & Serha, O.I. (2010). Klasyfikatsiia stepovoi roslynnosti Lisostepu Ukrainy [Classification of steppe vegetation of the Forest Steppe of Ukraine]. Naukovyi visnyk Uzhhorodskoho universytetu. Seriia Biolohiia, 27, 68–79. (in Ukrainian).
100. Yaroshenko, P. D. (1961). Geobotanika: Osnovnyie ponyatiya, napravleniya i metody [Geobotany: Basic concepts, directions and methods]. Akad. nauk SSSR. Izd-vo Akad. nauk SSSR, 138 (in Russian).
101. Zoz, I. H. (1933). Roslynnist Mykhailivskoi tsilyny na Sumshchyni [Vegetation of Mykhailivska virgin land in Sumy Oblast]. Zhurnal bio.-botan. tsyklu VUAN, 5–6, 157–183 (in Ukrainian).
Published
2023-01-20
How to Cite
Koplyk, Y. (2023). STEPPE PHYTOCOENOSES OF UKRAINE: SPESIFICS AND HISTORY OF THE STUDY. Bulletin of Sumy National Agrarian University. The Series: Agronomy and Biology, 49(3), 40-48. https://doi.org/10.32845/agrobio.2022.3.6